DNA recovered from an Athenian mass grave pointed to which agent for the Plague of Athens?

1. The Plague of Athens in 430 B.C. killed a quarter of the Athenian troops in the Peloponnesian War. 2. Researchers used DNA from teeth recovered from a mass grave. 3. They identified nucleotide sequences from Salmonella enterica serovar Typhi, which causes typhoid fever. 4. Typhoid spreads through feces and is common in overcrowded areas, which fits crowded wartime Athens. 5. Yersinia pestis caused the later bubonic plagues, not this outbreak. _Source: OpenStax Biology (1st ed., CC BY 4.0), Ch 22 "Prokaryotes: Bacteria and Archaea", section 22.4 Bacterial Diseases in Humans_